Table 1

Locomotor and behavioral variables

	Variable	Units
	Locomotor Variables
1	Radial velocity with respect to operant lever (mean)	mm/s
2	Radial velocity with respect to operant lever (maximum)	mm/s
3	Radial velocity with respect to operant lever (SD)	mm/s
4	Speed (mean)	mm/s
5	Speed (SD)	mm/s
6	Speed (maximum)	mm/s
7	Move duration	s
8	Path length	mm
9	Latency to time of maximum speed	s
10	Latency to time of maximum acceleration	s
11	Maximum distance between actual path and best (straight line) path	mm
12	Turn efficiency (cumulative summed change in heading/net change in heading)	None
13	Angular velocity with respect to operant lever (SD)	Radians/s
14	Angular velocity with respect to operant lever (maximum)	Radians/s
15	Angular velocity with respect to operant lever (mean)	Radians/s
16	Signed change in head orientation	Radians
17	Radial velocity with respect to operant lever (mean)	mm/s
18	Path efficiency	None
	Variables measured at moment of cue onset
19	The distance from the lever at movement-onset	mm
20	Time elapsed since last reward received	s
21	Time elapsed since last lever press	s
22	Time elapsed since last cue (DS or NS)	s
23	Head orientation with respect to lever	Radians